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The "author-pays" model allows PLOS journals to provide all articles to everybody for free (i.e., open access) immediately after publication.
Both numbers for 2016 roughly correspond to the industry-wide averages for biology-related journals.
In mid-2012, as part of a rebranding of PLo S as PLOS, the journal changed its name to PLOS One.
At PLOS One, the median review time has grown from 37 days to 125 days over the first ten years of operation, according to Himmelstein's analysis, done for Nature.
Reuse of articles is subject to a Creative Commons Attribution License, version 2.5.
In the first four years following launch, it made use of over 40,000 external peer reviewers.